INVESTIGATIONS INTO DRY-SEASON TOMATO PRODUCTION IN TRINIDAD. (With a general review of the literature on tomato cultivation.) by W. F. K. CHRISTIAN, B. Sc. (Hons. Bot.) D.T.A . Report A report submitted in part requirement for the Diploma in Tropical Agriculture of the Imperial College of Tropical Agriculture, Trinidad, T.W.I. JUNE, 1958. C O N T E N T S . PAGE. INTRODUCTION 1 REVIEW OF LITERATURE. 4 1. Origin of the cultivated tomato 4 2. Botany 5 3. Climatic conditions of growth. 6 (i) Temperature. 7 (ii) Light intensity and Optimal Night and Day Temperature 8 (iii) Light Intensity and Ascorbic Acid content of fruit. 8 (iv) Rainfall 8 4. Soil conditions. 9 5. Fertilizers 9 6. Starter solution 12 7. Training and Pruning. 14 8. Propagation and transplanting 16 9. Methods of Cultivation 19 10. Spacing in the Field 19 11. Varieites and Horticultural Characters of tomatoes 20 12. Hybrid vigour. 23 13* Diseases and Pests. 23 14. Fungal and bacterial diseases 23 (i) Fusarium wilt 23 (ii) Bacterial wilt 24 (iii) Septoria leaf spot or blight 25 (iv) Late blight. 25 (v) Stemphylium or Gray leaf Spot. 26 (vi) Early blight 26 (vii) Tomato leaf mould. 27 15. Virus Diseases 27 (i) Tomato (tobadco) mosaic 28 (ii) Cucumber mosaic 28 (iii) Tomato spotted wilt, (iv) Single or Tomato Streak,, (v) Double or severe tomato Streak, (vi) Curly top (vii) Control of Virus diseases 16. Fruit rots. Control of fruit rots. 17. Physiological diseases. (i) Blossom-end rot (ii) Growth cracks (iii) Puffiness (iv) Sunscald 18. Pests, (i) Ants. (ii) Millipedes (iii) Caterpillars, cutworms, fruitworms. (iv) Eelworms (v) Mites (vi) Aphids (vii) Leaf miner (viii) Midge 19. Grading 20. Packing for the market. 21. Storage EXPERIMENT. 1, Object of Experiment. 2, Experimental details. 3, Soil type 4, Nursery Operations. 5, Field Operations. (i) Transplanting. (ii) Moulding, weeding, fertilizing (iii) Pruning (iv) Staking Page* (v) Spraying 46 (vi) Irrigation. 4-7 6, General Observations in the Field, 47 (i) Weed growth. 4-7 (ii) Diseases 4-7 7. Harvesting 49 RESULTS 50 ANALYSIS OF VARIANCE. 53 DISCUSSION 56 SUMMARY 60 ACKNOWLEDGEMENT 61 APPENDICES ( I - IV) 62 BIBLIOGRAPHY. 68 I N T R O D U C T I O N . The tomato is one of the most popular as well as one of the most important vegetables in the tropics and temperate region. Rich in vitamin C and minerals especially phosphonis, potassium and calcium, and high in palatibility it is esteemed in various dishes0 In the tropics its increased consumption will go a long way in decreasing the many deficiency diseases found there. For most countries it is an important conmercial crop bringing in much needed revenue. Though a native of tropical Central America, the tomato has reached its maxinum production in temperate countries. In Britain the green house culture of the tomato new produces up to 60 tons per acre, the average yield being 35.0 tons per acre. This is about 2-g- times larger than the average yield of the outdoor crop produced in the U.S.A. In the U.S.A. the tomato ranks among the vegetable crops second only to the potato in farm value, and heads the list in value among the perishable vegetables. The average annual production and farm value for the crop grown for the fresh market for the period 1948-52 were 33,322 million bushels and 5133,893 million respectively. For the same period the production and value of the crop for processing were 3,144,700 tons and 588,398 mil lion (Thompson and Kelly, 1957). Fresh tomatoes in South Africa brought to the farmers during 1952-53 £1,310,000 on the nine principal markets of the Union. In Cuba and some of the islands in the Caribbean, principally Jamaica and the Bahamas, tomatoes are grown mainly for export to the U.S.A. during the winter and early spring when field production is not possible in most parts of the latter country. Montserrat ships its produce to Eastern Caribbean countries like Trinidad and British Guiana. Since the last war the former trade in fresh fruit between Canada and the British West Indian islands, Jamaica, Antigua, St. Kitts, Montserrat, Bahamas and Bermudas has stopped owing to lack of regular shipping. Other important tomato-grcwing countries are Mexico, Puerto Rico, the Canary Islands, and Hawaii. In../ 2 In the Canary Islands the crop is grcrvn mostly under irrigation and exported to Europe, the United Kingdom taking the greatest amount. Hawaii tomato cultivation has recently come very much to the fore and much scientific research has gone into it. The Mexican crop is exported to the U.S.A. In the tropics in general the tomato is grown mainly in the dry season, because high temperatures and continuous heavy rainfall during the wet season create conditions unsuitable for growth. Techniques of cultivation are backward and yields are low. Resistant varieties are not available and a large part of the crop is lost through diseases. Great improvement is therefore needed in the techniques of cultivation of this crop. For the past five years or more, based on the findings aft the John Iunes Horticultural Station that yields of tomatoes depend to some extent on methods of production of seedlings, experiments have been conducted at the I.C.T.A. into methods of raising seedlings with a view to improving on the existing local methods. Three methods have been compared: raising seedlings in 3" clay pots containing specially prepared composts; raising seedlings by the methods of the peasant farmers who sow nursery beds thickly with seed and do not thin out; and producing seedlings by the "improved" method at a spacing of 3" x 3" on open beds. The results have so far indicated that pot raised and "improved" seedlings are superior in yielding capacity than the "peasant" seedlings with the pot raised seedlings giving higher yields than the "improved" seedlings. Cultivation methods have also been investigated. Plants grown on ridges, on the flat and on the flat with subsequent moulding up have been compared. In most of the experiments ridged and moulded plants have out-yielded those grown on the flat. But there are as yet no clear cut, confirmed differences between ridged and moulded plants. The effect of Starter solution used in transplanting has also been probed into and found beneficial. In the experiment reported in this paper the intention was to confirm the effects on yields of the three cultural methods - planting on the flat, moulding and ridging - and also of pruning and staking. It was decided ../ - 3 It was decided in raising the Seedlings and in transplanting to use methods which were within the means of the peasant farmer and were capable of giving high yields<, The seedlings were therefore raised at a spacing of 3" x 3" straight on open beds instead of in pots or in seed boxes prior to transplanting to a field nursery, and a starter solution was used during transplanting. 4 REVIEW OF LITERATURE, Origin of the cultivated tomato* The centre of origin of the genus Lycopersicon, to which the tomato belongs, is on the narrow west coast area of South America, between the Andes and the ocean, extending from the equator to about 30° South latitude, Where the cultivated forms of the genus evolved in this area is however a matter of dispute. Though it is widely believed that the centre of origin of the cultivated forms was in Peru, Jenkins (1948) adduces evidence to shew that Mexico was the centre of domestication of the tomato, Jenkins points out that in Mexico there is a widespread distribution, not found in Peru, of both wild and cultivated tomatoes and of many transitional forms between the wild and cultivated types and also between the cultivated types themselves. He also draws attention to the fact that the name tomato is derived from the Nahua word "tomatl" which in Mexico is applied to many different solanaceous plants including lycopersicon, Saracha and Physalis especially the latter which was and still is used as food in many parts of that country. Though the tomato had been called many names including the European one of "Pomi del Peru" Jenkins, in his investigation^ found nothing in the historical records to suggest a, Peruvian origin. Prom the results of his researches Jenkins has put forward the hypothesis that the putative variety cerasiforme of lycopersicon esculentum was the ancestor of the cultivated tomato. Originally confined to the Peru-Ecuador area, this variety spread in pre Columbian times as a weed throughout much of tropical America. In mexico owing to its general resemblance to the older food plant Physalis it was brought into cultivation. Prom Mexico the cultivated forms were taken to Europe soon after the Conquest by Cortez in 1534, In Europe the tomato was at first probably grown in the gardens of herbalists as a curious plant but soon found its way into potions and later became a relished article of food. According to Sturtevant (1919) and McCue (l952) the earliest mention of its use as sauce in Italy was by Matthiolus in 1544 in the first edition of his commentary on Dioscorides, 5 Before 1600, however, it was widely established as a vegetable in many European countries. From Europe the Spanish and the Portuguese took the tomato to Asia, the Pacific and Africa (Jenkins, 1948). As regards the introduction of the crop into the United States of America, Thompson and Kelly (1957) record that the first reference to the vise of tomato for culinary purposes in the United States was by Jefferson in 1781, although mention had been made of its presence in the Carolinas, Georgia and Florida earlier in the 18th Century. Botany. The tomato, botanically known as Lycopersicon esculentum Mill, belongs to the family Solanacea (L). Cobley (1956) has given a botanical description of the plant, which is an annual or short lived perennial with weak trailing much branched stems covered with yellow glandular hairs as well as non-glandular ones. The fruit is a two - to- many - celled red or yellowish berry with a smooth skin. The seeds are many, small, hairy, kidney shaped and are borne on fleshy placentae in axile placentation. The red colouration of the fruit is dus to the presence of two pigments carotene and lycopersicon occurring in different concentrations and thus bringing about gradation in fruit colour. Both Muller (1940) and Luckwill (1943) have divided the genus Lycopersicon into two subgeneraj first, Eulycopersicon including the red fruited species L. esculentum and L. Pimpinelli folium; second, Eriopersicon, the green fruited species which according to Muller consist of the Species L. cheesmanii, L. peruvianum, L. hirsutum, and L. glandulosum. The red fruited group including all cultivated forms has a complex geographic distribution that has been greatly influenced by man. 6 The green fruited group contains no species of economic importance. L. pimpinelli folium, one of the two red-fruited species has a restricted range of distribution being confined to Peru, Ecuador, and the Galapagos Islands (MuLler, 1940). The variety Cerasiforme and the form pyriforme are considered by Muller as distinct entities within the species esculentum. Tomatoes may also be divided into determinate and indeterminate types according to the sequence of formation of flower clusters. Shoemaker (1949) has given the characteristics of the two types. Indeterminate plants are capable of continuing their growth more or less indefinitely. Normally a blossom cluster is produced at every 3rd internode, being separated by 3 leaves from the next flower cluster. Determinate varieties are free blooming. The blossom clusters are produced at every internode, thus being separated by only one leaf. The stem eventually terminates in a blossom cluster. Climatic Conditions of Growth. The tomato is mainly a crop for warm conditions, but may be grown in frost-free areas during the winter in temperate regions. To give maximum yields it requires a long growing period of 3^ - 5 months with moderately high temperatures, (Van HLommestein et al, 1954). High tenperatures coupled with high humidity, however, favour various fungus diseases, while hot drying winds often result in the dropping of the blossoms, (Thompson & Kelly, 1957). The tomato plant therefore" produces largest crops in cool dry regions under irrigation and also under the controlled conditions that obtain in a glass house. 7 Temperature. Work by many investigators has shown that temperature has a marked effect on the growth and setting of fruit of the tomato. Watts (1931), found that fruit set was greater at 75° than at 60°F. Moore and Thomas (1952), also found out that when the average maximum day temperature was above 90°F and the average minimum night temperature above 70°F fruit set was low. Went (1945), demonstrated that night temperature was actually the critical factor in the setting of fruit by tomatoes. Fruit set was abundant only at night temperatures between 15° - 20°C under which conditions stem elongation was also optimal. Less growth and much less fruiting occurred at either higher or lower night temperatures. This temperature effect, Went observed was much greater than that of the relative humidity of the air, length of day or day temperature. Went and Cosper (1945), found that in the field the effective low temperatures usually occurred around 30 days before maximum fruit production i.e. they coincided with the period of fruit setting. The effect of temperature on fruit production and growth of tomatoes has been attributed by Went (1945), to the direct effect of temperature on size of inflorescence and trans­ location of Sugars. Went in his investigations discovered that the size of the inflorescence varied very much with night termeratures. The warmer the night, the smaller the inflorescence and the flowers. It was only at night temperatures between 13° C and 22° C that fruits started to grow. At higher temperatures no fruit was set at all. Above 18° C night temperature, translocation of Sugars became limiting to the growth processes of the whole tomato plant. In varieties that Went studied he found thermo-periodicity responses to be consistent and large enough to characterise them. 8 light Intensity and Optimal Night and Day Temperatures. •When tomato plants are exposed to lover light intensities there is a shift towards lower optimal day and night temperatures. (Went, 1945). Growth of plants at reduced light intensity and high night temperatures (26° - 30° C) was very small, while at low night temperatures reduced light had no effect. Hoffman in 1938 obtained similar results in Ohio with plants grown in a glass house during the spring. He found that the best day temperature for Globe tomatoes was 18° C on cloudy days and 21° - 24° C on Sunny days. light Intensity and Ascorbic Acid content of Fruit. Of the factors day length, light intensity, temperature, fertiliser, Soil type, and humidity, Hammer et al (1945), have shown that light intensity previous to harvest has the greatest influence on the ascorbic acid content of the fruit. Under low light intensities the ascorbic acid content is much lower than under high light intensities. Increases in the ripe fruit of 66# in ascorbic acid resulted when plants were transferred from shade to sunshine at the time the fruit was mature - green. Hammer and his co-workers suggest that much of the effect on ascorbic acid content ascribed to season and location may be due to variations in light intensity. Rainfall. This is important only for the out door crop. In Trinidad (Rombulow-Pearse, 1953) and generally throughout the tropics tomatoes are mainly grown during the dry season usually with irrigation. Excessive rain increases the incidence of disease, destroys newly formed blossoms and according to Young and Mac Arthur (1947), decreases pollination to cause puffed fruits, which contain air spaces in locules incompletely filled 9 with seeds. Soil Conditions• Many types of soils ranging from sands to clays have been found suitable for the cultivation of tomatoes. 66$ of the entire crop of Peninsula Florida is gram on sandy soils vhich are either acid or caleareous, (Spencer, et al. 1955). Hardy and Baker (1952) found tomatoes growing satisfactorily on the highly calcareous black, marly rendzinas of the swash-land in Andros Island. Loamy soils are however preferred for growing tomatoes. It is agreed by most workers that a well-drained soil is essential for high production. A gradual slope facilitates irrigation and in general provides excellent drainage. Van HLommestein et al, (1954), state that provided the drainage is good the depth of soil below 18" is of less importance. The tomato has been said to be tolerant of acid soil conditions. Excessively acid soils should however be limed to reduce acidity and create more favourable conditions for the soil bacteria and decomposing organic matter. Sayre (1947), reported an increase of 4.2 tons of fruit per acre from an application of 3 tons of dolomitic limestone to the acre on Fulton loam soil of pH 4.8. Spencer et al (1955) recommended a pH value between 5.5 and 6.0 for tana toes grown on sandy soils, and at pH values above 6.0 advised no liming but a constant watch for possible trace element deficiency symptoms. Fertilizers• Although animal manures are used to fertilise the soil for tomatoes, artificial fertilizers, especially nitrogen, play a greater role in tomato culture. 10 Hie amount of nitrogen per acre used by many growers has however varied betvreen very wide limits, and the same amount of nitrogen used in different areas has also been known to give different results. It has also been shown by Kraus and Kraybill (I9I8), that over fertilisation with nitrogen causes the plant to be vegetative and produce little fruit. In a review of soil practices with some outstanding crops in New Jersey, Hester (1940), observed that the amount of Nitrogen used by certain growers varied from practically none to 140 pounds per acre and also that in certain c ases where different growers used an equal amount of nitrogen, namely 75 lbs. per acre, a tremendous variation in yield from less than 5 tons to nearly 15 tons per acre was obtained. While other factors influence the yield of tomatoes, the indication of the above results was that response to fertilizer might depend on the time and method of application. Hester (1938), and Hester et al (1951) doing chemical analyses of tomato plants at monthly intervals showed that a large portion of nutrients is absorbed by the plants between the 2nd and 3rd months after transplanting in the field and that the nutrients absorbed in the largest quantities are potash, nitrogen, and calcium. Hester considers it desirable, therefore, to apply part of the fertilizer before the plants are set out and the remaining part later, Hester (1941), found on Sassafras fine sandy loam that the use of nitrogen and potash in two side dressings was more efficient than either mixing the fertilizer in the soil in the rcw before planting or broadcasting and ploughing down later, Hester however quotes the unpublished results of experiments in the middle West States as giving indication that broadcast application and ploughing down on heavy soils of various sources of nitrogen as being the most efficient of all methods tried. Jones and Warren (1954) also compared various methods of placement of phosphorus and found that deep placement under the row was more efficient than banding beside the plants or broadcasting. 11 Though opinion on the most beneficial feitilizer to the tomato crop is somewhat divided many experiments have indicated that depending on soil type, nitrogen and phosphorus would appear to be the most important fertilizers to the crop and potash probably the least since the element is often not very much lacking in soils. The use of 100 pounds nitrogen per acre on a Coto clay soil in Puerto Rico gave significantly higher yields of commercial tomatoes per acre, than either none or 200 pounds nitrogen per acre, (Landrau and Samuel, 1955). Increased yields were also obtained on the same soil with phosphate fertilizers, the highest yields beiig obtained with 200 pounds of Pg O5 per acre. There was no response to potash on this soil. Miller (1954) working on the effects of various fertilizer mixtures on the yield and growth of tomatoes found that a couplete mixture of NPK used at 14 cwts. per acre produced the highest yields and the best vegetative growth. A f ertilizer mixture containing only phosphorus and nitrogen had about the same effect as the couplete mixture, while a mixture containing phosphorus and potassium gave the most unthrifty growth and the lowest yields. Miller observed that the absence of nitrogen in a fertilizer mixture caused a marked reduction in grcwth rate while a coirplete mixture containing potassium lengthened the period of growth. Chin (1932) found super phosphate and sulphate of potash at 200 lb. per acre depressed yields of the Marglobe variety while Ammonium Sulphate at 200 lb. per acre definitely increased yields by about 1.2 tons. Results of long continued experiments in Pennsylvania conducted by Warren et al, (1940) also shewed that the greatest yields were always obtained using a complete fertilizer and that as the phosphorus content of the fertilizer increased, yields also increased. Fhosphorus it was found was more effective in increasing yields than either nitrogen or potassium. Organic manure from various sources has been found on many occasions to increase yields of tomatoes. In the Pennsylvania experiments (Warren et al, 1940 ) 20 - 40 tons of well rotted farm yard manure were found superior to any commercial fertilizer mixture tried. 12 In Puerto Rico Lardrau et al (1955) obtained significant increases in yields when they used fitter press cake, and organic product of the Sugar mills averaging about 2,5# Na and Pg 05 at 20 tons per acre. When combined with 100 pounds of nitrogen and 200 pounds of Pg Og per acre the yields obtained were higher than those obtained when the fitter press cake or chemical fertilizer was used alone. Cotton seed meal has also been found effective as fertilizer in Montserrat, (Campbell, 1957). Dried blood is also sometimes used as fertilizer especially in green house tomato culture (Allerton, 1954). On Andros Island, Bahamas, Hardy and Baker (1952) found "MUorganite" an American Sewage product containing 6-8% Nitrogen and Z - 6% a vailable phosphate being used with success in combination with inorganic fertilizers• Fertilizer mixtures recommended for use on tomatoes vary in composition according to soil type. Thompson and Kelly (1957) recommerd It 2: 1 and lj 2; 2 mixtures for use on lighter soils where tomatoes are grown in rotation with other cultivated crops. For loam, silt loam, and clay loam soils where also the crop is grown in dotation with field crops they recommend mixtures of It 3: 1 and lj 4: 1. The Florida Agricultural Service recommends for lighter soils a ratio of 4: 7j 5 and 4: 8: 8. Patterson (1932) suggests a mixture of one part nitrate of soda to two parts of superphosphate and one part of muriate of potash applied at the rate of 500 lbs per acre in field dressing before transplanting. A f urther 600 lbs should be applied later as side dressing. Starter Solution. In the United States and in Europe it has long been a common practice among growers to transplant tomatoes with from \ to 1 pint of solution of sulphate of ammonia or nitrate of soda in water. Baker working in Northern Indiana in 1937 reported increased yields of tomatoes and earlier ripening where commercial phosphoric acid and mono- ammonium phosphate were used in the transplanting water. Grace (1937) reported better results from very minute quantities of hormones in the transplanting water and stated that with tomato seedlings increased growth was obtained with naphthylacetic acid at a rate corresponding to 200 mLlli grams per acre. 13 Sayre (1938) confirmed Baker's results and obtained significant increases in yields with Ammonium phosphate and nitrate of potash in the transplanting water. He also found that the mono-ammonium phosphate was a significantly more effective form of phosphorus than the ortho-phosphoric acid in stimulatirig plant growth; also that the use of small amounts of borax, manganese sulphate and magnesium sulphate in the transplanting water did not increase yields. As regards the effect of hormones on growth of tomatoes Sayre found contrary to the results of Grace that hormone had a depressing effect on yields, particularly the early yields. In later investigations Sayre (1939) found that a mixture of two parts of Ammonium phosphate and one part nitrate of potash was as effective and less expensive than any other combinations of chemicals he tried, Sayre also discovered that increasing the concentration and reducing the amount of solution per plant increased the effectiveness of the nutrient solution. These findings have been corroborated by work done by Tremeer (1956) and Fenwicfc (1957) at the Imperial College of Tropical Agriculture. These workers obtained increased growth rate, early flowering and increased yields with starter solutions containing 2 parts of Ammonium phosphate and one part nitrate of potash. Fenwich comparing two levels of Starter Solution also found that, the solution applied at the rate of J pint per plant in a more concentrated form was more effective in increasing yields than that applied diluted at \ pint per plant. Porte (1952) however finds the use of starter solution unnecessary where fertilizer has been placed in the rows before planting. The principal effect of a Starter Solution used in transplanting tomatoes is to enable the plant to become established quickly. This results in a marked gain in early maturity. Sayre (1939) was of 1he opinion that the entire nutrient solution being in immediate contact with the roots and in a concentration sufficient to saturate the fixing capacity of the soil, was readily available to the roots just at the critical time of transplanting. A s mall amount of concentrated solution was more effective because it resulted in less puddling of the soil around the plant with less cracking as the soil dried. 14 Training and Pruning, In many tomato growing countries training and pruning tomatoes is a common practice. Various methods of training are employed. The pole or stake method is however the commonest used both for the green house and outdoor crops. In this system a strong stake 5-7 feet high is set securely alongside each plant soon after the first blossoms appear. The plant is tied to the stake with soft twine (Thompson and Kelly, 1957), rag string, hemp or raffia (Hieronymus, 1949) Allerton (1954), looped around the plant and tied around the stake. The tying is continued up the stake as the plant grows taller. Where the crop is gram in double rows the stakes may be slanted to meet in wigwam formation at the top where a strong tie is made, (Allerton 1954), This method is particularly suitable for exposed fields which cannot be adequately sheltered artifically. The plants instead of stakes may be supported by heavy twine running from the base of the plant to an overhead attachment. Pruning is done by removing either by hand or with a knife the new s hoots that appear in the leaf axils once every week or ten days. The plant is most commonly pruned to a single stem but there are growers who prefer to have two or three stem plants. Other methods of trellising are nearly always more elaborate than the stake method described above. In Hawaii the determinate varieties which trail on the ground are supported on low racks 12-15 ins. high at the back and sloping to ground level towards the plants, (Hieronymus, 1949), In South Africa several methods are in use. In one method described by Van HLommestein, (1954) 2-4 Spanish reeds or stakes are erected in a tripod formation around each plant. In another method, also described by Van Blommestein, 2-4 wires one above the other are stretched along light stakes and plants tied to them. In a modification of this method pairs of parallel wires are stretched so that the plants are kept upright between them. This method apparently is also used in New Zealand (Atkinson et al, 1949), 15 Thou# there seems to be no agreement as to the value of staking and pruning, the following advantages as summarised by Deorrieret al (1944) are usually claimed for this treatment: (1) Earlier ripening. (2) Larger fruits. (5) Less disease injury. (4) Larger early yields. (5) Cleaner fruit. (6) More convenient harvesting. (7) Greater convenience for spraying The disadvantages usually set against the advantages are, (Deonier et al 1944) (1) Greater amount of labour and expense, (2) Less total yield per acre, (3) Greater loss from blossom end rot, (4) More sunburn injury to the fruit, (5) Greater number of cracked fruits. The apparent conflicting results reported by many workers were probably due mainly to the use of less space or more plants per acre for the pruning and staking practices as compared to the untreated plots, and probably also to the wide variations in environmental conditions under which the different trials were conducted. From the results of many experiments, however, it would appear that pruning and staking increase early yields but bring about reduction in total yields. (Thompson, 1934j Hawthorne, 1939; Currence, 1941; Magruder, 1924; Walker, 1927). Thnmpson (1934) and Hawthorne (1939) attributed the increase in early yields caused by pruning and staking to the increased number of pruned plants per acre, the result of the closer spacing usually adopted for this treatment. Topper, however, in 1942 obtained at I.C.T.A. greater yields for staked and pruned plants when compared with unstaked and unpruned plants, at the same spacings, these being 2* x 3', if' x 3', and V x 3' • Results of experiments conducted in Mississippi and reported by Deonier et al (1944) also indicate a decided advantage for staking with pruning, and no advantage for staking without pruning over no staking and no pruning. 16 Riollano (1944), however, working in Puerto Rico found staking reduced consistently and significantly yields of all grades of fruit when compared with unstated treatments. Staking however improved percentage of marketable fruit but this was offset by the marked reduction in yield of marketable fruit. The combination of staking and pruning, Riollano found, reduced yields further in a significant way. Styrdom (1955) in South Aftica also found staking with pruning to have on yields adverse effects which were more noticeable at closer spacings. Pruning also retarded ripening and did not produce an earlier crop. Conflicting results of pruning and staking may also follow, according to Currence (1941), when different varieties of tomatoes are subjected to pruning and training. Currence found that pruning was beneficial to the Break-O'Day variety and detrimental to the determinate Pritchard. The fact that the incidence of the two physiological diseases,blossom end rot and fruit cracking is often more severe on pruned trained plants and also under dry and wet conditions indicates that pruned, trained plants either enable the soil to dry out more quickly or are unable to use soil moisture as well as unpruned plants. Studies by Thompson (1927) suggest that both these factors are probably involved. Propagation and transplanting. Tomato seedlings for transplanting can be raised by one of two methods. For the green house crop the seedlings are raised in soil blocks or clay, paper or plastic pots filled with specially composted soils. The seedlings are later transplanted to the field with a ball of soil. In the second method which is mostly adopted in the U.S.A. and in the tropics, seeds are raised on open beds and later transplanted with bare roots to the field. In the tropics the peasant farmer sows seeds thickly on the open bed and usually fails to thin out, thus producing seedlings which are over crowded, have sparse foliage and roots , and look spindly and weak. The methods for the propagation of seedlings in the U.S.A. have been outlined by Porte (1952) . In areas where the growing season is long seedlings are raised in cold frames or open beds, at a spacing of 12" - 15" between rows and 1" - 2" between plants in the row. Where the growing season is short and where also earliness is important the seeds are sown in 17 green hotees or hotbeds and the seedlings are later transplanted to a wider spacing in hot beds or in cold frames. Direct seeding in the field is also carried out in the U.S.A. Hie seed is machine drilled in rows 15" - 18" apart at a rate which gives 15 - 18 plants per foot of row. Hiese are later thinned out to the required spacing. This method is applicable only in places like California, Nevada and parts of Florida where frost is not a hazard and normal soil temperatures are suitable. It has the advantage of saving labour, of transplanting and often reduces loss from damping off. In Trinidad and possibly in other parts of the Tropics, direct seeding may prove a complete failure. Rombulow- Pearse (1953) in Experiments at the I.C.T.A. found it difficult to establish plots by directly seeding in the field because the soil tended to bake hard and ants carried the seeds away. Where direct seeding in the field has not been done tomato seedlings raised in pots or on open beds are eventually transplanted into the field. Experimental results reported by Loomis (1925) indicate that transplanting does not directly promote the growth of the plant and is of no particular agronomic value, but may be an economic expedient in saving valuable green house or garden space and in allowing for better care of slowly growing seedlings. In the early seedling stage tomato seedlings can be transplanted with little or no injury. With increasing maturity hcwever the danger of injury increases. This is in agreement with the results of Casseres (1947), (Babb 1940), Brasher and Westover (1937) and Porter (1935) which show that young plants that are tender and capable of quick growth resumption after field setting give early and large total yields. Any treatment of the seedlings which results in hardening reduces the early and total yields in proportion to the degree of hardening. In this connection Casseres for plants grown at 60 - 70°F ohtained results in which 7 week old seedlings significantly produced higher early and and total yields than seedlings 11 weeks old. Under tte warmer conditions in Trinidad 4 weeks old seedlings are considered ideal for transplanting (Rombulow-Pearse, 1953; Barett I955j Hremeer, 1956; Fenwick, 1957). Yields of tomatoes are known to depend very much on the way the seedlings are raised and the treatment given to them. 18 Working at the John Lunes Horticultural Institute, Lawrence (1950) was able to show that the quality of compost used in the pots, the time of pinching out young seedlings from seed boxes to the pots, the size of the pot and hence the spacing between plants, all affected early and total yields. Work and Amy (1950) found that plants grown in beds and thinned to a spacing of 2" x 2" or 3 or 4 seedlings per inch gave significantly higher yields than plants from overcrowded beds. Similar results were obtained by Smitii (1954), Barett (1955), Treemeer (1956), Fenwich (1957) working at I.C.T.A. fliey found that "improved" seedlings i.e. seedlings raised on the open bed at a spacing of 3" x 3" gave higher yields and necessitated less supplying than the overcrowded seedlings raised by the methods of the peasant farmer. Pot raised seedlings are considered superior to seedlings produced in the open bed. Nylund (1956) reported results in which transplants grown in 3-inch pots outyielded those grown on flat open beds at 2" x 2". Snyder showed that seedlings in pots of this size may even outyield seedlings grown at a spacing of 4" x 4" on flat beds. Experiments at I.C.T.A. have also shown that seedlings raised in pots yield better than those produced on beds (Treemeer 1956, Fenwick 1957). Bie increased yields and better field establishment given by pot raised seedlings and seedlings produced at a wide spacing on open beds have been attributed to better root and foliage development. At the I.C.T.A. Smith (1954) Treemeer, (1956), Fenwick (1957) found that pot raised and "improved" seedlings had well developed roots and leaves while seedlings raised on overcrowded beds had hardly any roots and leaves. Loomis (1925) also considered the high yields obtained from seedlings raised in pots to be due to the well developed roots and luxuriant foliage of the seedlings. I When these well developed seedlings are set in the field they are less subject to mechanical injury, to disease and insect attack, and to destruction through drought. 19 Methods of Cultivation. Field tomatoes may be transplanted on ridges or on the level and earthed up later when found necessary. In Puerto Rico level or flat planting with irrigation is favoured during the dry season while on heavy soils during the rainy season ridge planting is considered best, (Childers et al 1950.) In Ceylon planting on ridges in wet districts and on the flat with subsequent moulding up in dry areas is also practised, (Senaratire and Senathiraja, 1927). In the Canaries tomatoes are grown only on ridges, (Holmes, 1931), In Trinidad the peasant farmer who grows tomatoes mainly in the dry season xisually plants on the level and earths up later. (Rumbulow-Pearse 1953). Experiments at the I.C.T.A. have demonstrated that plants grown on ridges and on the flat with subsequent moulding up give higher yields than those planted on the level and not earthed up, (Barett, 1955; Fenwick 1957; Smith, 1954). The spacing given to plants in the field varies greatly according to variety grown, methods of cultivation, and whether the plants are pruned and staked or allowed to trail over the soil. Many experiments conducted on field spacing have, however, shown that close spacing results in larger yields than wider spacing. Currence (1941) working in Minnesota reported results in which closer spacings uniformly increased yields which reached a maximum at a spacing of 1' x 4'. There was a tendency to smaller fruits at the closer spacings but the tendency was not uniform and definite enough to show significance. Reeve and Schmidt (1952) also obtained higher total yields from plants given 7 square feet of space than those given greater space. Similar results have been obtained in St. Kitts, and Antigua. In St. Kitts (Anon. 1931) a spacing of 4* x 2» gave yields significantly better than 4' x 3», and in Antigua ( Anon. 1949) higher yields were obtained at a spacing of 3* x 1' than at 3' x 2'. 20 Large-growing spreading varieties require more room than small- growing ones. Spacing for the spreading varieties should however not be so large as to reduce yields. Where large mechanical equipment is used for spraying and dusting wide spacing is also required. In the U.S.A. a compromise is reached by having rows 4- - 5 or more feet apart and setting the plants close in the rows. (Thompson and Kelly, 1957). Staked and pruned plants are usually planted at closer spacing than unstaked and unpruned plaits. (See also training and pruning). Varieties a^d Horticultural Gh racters of Tomatoes. Tomato exists in many distinct varieties and in numerous strains, and selections of the varieties. The choice of any particular variety for growing depends on soil type, climate, length of growing season of the area or whether an early crop is desired or whether the crop is being produced for processing. The ideal variety for any region should, however, combine most of the desirable characters including prolific fruiting habit, production ! 11 1 1 S R Q R 58.75 64.75 in. _1 : P M [ Q M 71.50 61.00 BLOCK III. , j BLOCK IV. 51 INDIVIDUAL PLOT YIELD OF FRUIT OVER 2" IN DIAMETER IN LBS. BLOCK I. BLOCK II. BLOCK III. BLOCK IV. S F 30.00 P M 22.25 I P M 39.25 i i P R 27.75 S M 25.00 P F 31.25 Q F 35.25 Q M 21.75 Q.M 33.25 S R 38.25 Q R 41.25 • | Q R 19.00 P F 39.50 S F 29.25 S R 43.25 S F 18.25 P R | ] 43.50 i , Q F 29.00 P R 40.00 S R 22.25 | I Q R 38.50 P R 38.75 P F 47.75 P F 15.00 Q F 35.00 S M 41.25 S F 39.25 S M 16.00 S R 33.00 Q R 38.75 Q M 36.75 JO o o VJX o P M 39.00 Q M 36.50 S M 40.00 P M 23.75 » ' 52 TWO - WAY TABLES. I. TOTAL YIELD OF FRUTT IN LB. r FLAT ; MOULD I RIDGE TOTAL I MEAN STAKING WITH PRUNING 201.00 224.. 50 230.75 656.25 54.63 STAKING WITHOUT PRUNING 230.00 219.25 255.00 704.25 58.69 NO STAKING; No PRUNING. 195.25 220.00 233.25 643.50 54.04 TOTAL 626.25 663.75 719.00 2009.00 MEAN 52.19 55.31 59.92 55.31 II. TOTAL YIELD OF FRUIT OVER 2" IN DIAMETER IN LB. FLAT MOULD RIDGE L . . _ TOTAL MEAN STAKING WITH PRUNING 116.75 122.25 i 136.75 375.75 31.31 STAKING WITHOUT PRUNING. 133.50 124.25 150.00 407.75 33.98 NO STAKING; NO PRUNING. 115.75 128.25 K I II 137.50 381.50 31.79 TOTAL 366.00 374.75 424.25 1165.00 MEAN 30.50 31.23 j 35.35 32.36 ANALYSIS OF VARIANCE (a) TOTAL YIELD. Source of Variation. D F S S M S I F Blocks 3 4305.24 1435.08 32.63 * * * Treatments 8 629.26 78.66 Cultural 2 362.82 181.41 4.12 * Training & Pruning 2 152.00 76.00 1.73 Interaction 4 1U.44 28.61 0.65 Error 24 1055.51 43.98 TOTAL 35 5990.01 S. E. of a single plot yield Coefficient of variation of single plot yield S. E. of mean of 12 plot yields = 6063 = 11.8 % 1.916. Training and pruning treatments and their interaction with the cultural treatments were not significant. The cultural treatments were significant at the 5% l evel. The F-ratio for blocks was very highly significant suggesting that there were real fertility or edaphic variations in the area and thus justifying the adoption of the randomised block design for the experiment. This fertility trend in the area was actually observed by its effects throughout the experiment. It appeared soil conditions were least favourable on block IV. which gave the lowest yield and took the greatest number of supplies. On this block also it was observed that the condition of the plants was not as satisfactory as it was on the other blocks in the early stages of growth. Applying the T - test to the cultural treatments it was found that at 24 D - F. ridging gave significantly greater yields at the 1$ level than planting on the flat. The increased yield of ridging treatment over the moulding treatment was however not significant. The difference in yield between moulding and planting on the flat was also not significant. Value of t for diff. between means of Ridge and flat = 2.877 ** it w n n n it n ti Ridge and mould = 1.715 it ii « n ii ii ti ii Mould and flat = 1.116 (b) FRUIT SIZE. Source of Variation D F S S M S i F Blocks 3 2031.06 6 77.02 *#* 28.70 Treatments 8 246.52 30.82 Cultural 2 164.44 82.22 3.48 * Training & Pruning 2 48.50 24.25 1.03 Interaction 4 33.58 8.39 0.36 Error 24 566.24 23.59 Total 35 2843.82 S.E. of a single plot yield Coefficient of variation of single plot yield S.E. of mean of 12 plot yields 4.85 15.055 1.4 lbs. 55 Training and pruning treatments and their interaction with the cultural treatments were not significant and do not seem to affect the size of fruit. The cultural treatments were however just significant at the 5% l evel. This suggests that fruit size may depend on the type of cultural treatment given to the plants in the field. "T" Test. Application of the "Trt test to the cultural treatments showed that at 24 D.F. plants grown on ridges yielded significantly at the 5% l evel larger fruits than either plants grown on the flat or moulded up. There was no significant difference between the flat and the mould as regards fruit size. Value of "t" for diff. between means of Ridge and flat - 2.449* n n it it n n ii ii Ridge and mould - 2.081* •» « " " « " " " Mould and flat - 0.36 56 Discussion, Cultural treatments. In the experiment plants grown on the ridges very- significant ly outyielded those planted on the level but showed no measurable difference from those planted on the level and moulded up later. No measurable difference in yield was also found between plants on the level and those on the "mould", though the latter outyielded the former. With the exception of Smith (1954-) who reported results in which flat cultivation and moulding significantly produced higher yields than ridging, various investigators at I.C.T.A. have obtained results similar to those obtained in the present experiment, in which plants on ridges and on the mould have outyielded plants on the level. Opinion is however divided on which of the two methods - ridge planting and planting on the mould - is superior. Fenwich (1957) obtained no significant differences between the two methods but had higher yields from plants on ridges, while Rombulow-Pearse (1953) also obtained no significant differences between the two methods but had higher yields from plants in the mould. The conflicting results may have resulted from the different conditions under which the experiments were done. Most of the experiments had an attack of diseases which in the case of Smith's work caused very serious damage; a fact which might well have accounted for the very different results he obtained. In some of the experiments also all the plants were staked to facilitate harvesting while in others staking was adopted as a main treatment whose effects were to be found out. Though no interactions have been reported between staking and cultural methods, there is evidence elsewhere that staking may interact with variety (Currence 194-1) and with spacing (Deonier, 1944)* / fr>0- too- ft . 1- 400- -'p- t p 6i 3 Qd U- LL 5oo- o o O ± ^K>- 3oo- £oo- JOO— G R A P H X ( S k o w t n ^ C U v a ^ t c l c c - i i v e " y v e A c i s ) £0*- cu|-fu,ra-C 'frect4u^ev«4"^ > • ' . ' F LAT IHouwD K»*>ci.e 1 r 19 V ~i r~ 2.5 2-7 F • E 13 HuA-P-l J >Axe oF T 3 €> f t a r c h T- «o H-ARVtFSTir^ In the present experiment it was found that plants on ridges produce larger fruits than those growing on the level or on moulds* This needs further confirmation as the level of significance is rather low. Training and pruning treatment^t Though the Training and pruning treatments showed no significant differences among themselves in yield, the indication would appear to be that staking and pruning do have adverse effects on early though not on total yields. Staking without pruning appears to retard early yields more effectively than the combination of staking and pruning. This increased early yield with staking and pruning is however nothing like as great as the early yields obtained with no staking and no pruning. (See Graph II). The higher total yields obtained for the two treatments staking with pruning and staking without pruning may be attributed to the increase in the incidence of disease after the first four pickings. After giving unsatisfactory early yields the staking without pruning treatment began to show increasing yields and over the last four pickings actually outyielded the other two treatments. The no-staking and no-pruning treatment was the hardest hit with disease and at the end of the experiment was showing an overall total yield below that of the other two treatments even though it had given the highest early yields. Under disease conditions it would appear staking is more advantageous than no-staking. This advantage may be lessened by pruning staked plants. The costings show that the most profitable treatment was staking without pruning, the next profitable no-staking and no-pruning and the least profitable staking with pruning (See appendix IV). Under conditions at the College farm, and from the stand-point of financial gain, staking with pruning is the least desirable of the three treatments. Despite the fact that it resulted in higher total yields than the no- staking and no-pruning treatment, it brought in a lower net income than 3oo- — STAKm<<;N«> ^ t a » < \ ( H Q A - k i j P f t u r ^ m q • • * Ho StakikG 5 Ko pew NiN(, lOo G R A P H ii (^Uovjomcj Cu^vuc^(ct4•v€ X~o~Y 'frUiyi\is\a atocl prMr)t*4 J4v^cc4i^Vs) 1 1 i—i— H 19 2.1 Fe. e RuA RY J>*te <5F i^AR VES ue\C, 1 € PI A R ch - 59 the latter treatment owing to the high cost of the staking and pruning operations. Pruning with staking may therefore not be recommended under any circumstances while staking alone may be recommended where there is disease risk or in the wet season. When, however, diseases are not a serious problem, tomatoes may be grown with the best results without staking and pruning. 60 S U M M A R Y . The effects on yields of training and pruning and of different cultural treatments are investigated. Planting on the ridge is found to give significantly higher yields than planting on the level, but to show no measurable difference from planting on the "mould". No measurable difference in yield is also found between planting on the level and planting on the "mould". No significant differences are found between the training and pruning treatments. There is indication, however, that growing tomatoes without staking and pruning favours early yields, and staking and pruning tend to increase total yields. The different operations in the experiment are costed for and it is shown that staking and pruning is of the least financial benefit, while staking without pruning brings in the greatest financial returns. It is however suggested that under conditions where disease is not a hazard tomatoes may be grown with maximum benefit without being staked and pruned. 61 A G K N O W L E D G E M E N T . The writer wishes to thank Mr. J. S. Campbell for his invaluable advice and guidance throughout the duration of the experiment. He would also like to express his thanks to Mr. G. E. Hodnett who advised on the statistical design and to Mr. M. Benny who supervised the field work. 1 62 A P P E N D I X I . Plan and Layout of the Experiment. Bed 8. Bed 7. Block I. 1 S F 2 S M 3 Q M 4 P F 5 P R 6 Q R 7 QF 8 S R 9 P M LEGEND. Q F = Planting on the flat; no pruning, no staking. Q H = Moulding; no pruning, no staking. Q R = Ridging; no pruning, no staking P F = Planting on the flat; staking no pruning. P M = Moulding; staking no pruning. P R = Ridging; staking no pruning. S F = Planting on the flat; staking and pruning. Block II, 10 I P M 11 P F 12 S R 13 S F 14 Q F 15 P R 16 S M 17 Q R 18 Q M Block III. 1 19 P M 20 Q F 21 Q R 22 S R 23 P R 24 P F 25 S F 26 Q M 27 S M Bed 9. B l o c k I v , - . 28 P R 29 Q M 30 Q R 31 S F 32 S R 33 P F 34 S M 35 0. F 36 P M w 135' 15 ft * i "1" S M = Moulding; staking and pruning. S R = Ridging; staking and pruning, N. - 63 - A P P E N D I X I I . c o S T S OF PR 0 D U C _T 1 0 N. Figures are derived from the records of 1/3 of an acre and converted to cost per acre. Operation. Post in Dollars (BWl). Raising of Seedlings. Preparation of nursery beds 8. A3 Labour - application of fertilizer. 3.06 Labour - building Shed and sowing. 25.29 Thinning 27.27 Labour - spraying perenox 2<,52 Cost of seed - A o zs. "Ogier" 14.#4-0 Cost of art. fertilizer - 30 lbs. 7.20 88.17 Field operations and transplanting. Marking blocks and plots 25.02 Making ridges 16.86 Ridging Holing and transplanting. 85.53 Cost of starter solution 7.29 Cost of aldrin - -J- g allon. 5«52 1A0.22 Operations during growth. Labour - spraying with perenox 36.03 Labour - spraying with manzate 30.06 Supplying. 9.27 Application of fertilizer. 25.02 Weeding and moulding 115.89 Labour - irrigation 8.37 Harvesting 185.67 Cost of Art. fertilizer - 330 lbs. 79.20 Cost of "perenox" 0.75 Cost of "manzate" - 2 lbs. 20.OA Cost of "fosferno" - 150 ec. 5.25 Cost of Dupont Spreader sticker - 32 ec. 2.AO 517.95 Total expenses for cultivated operations 7A6.34 64 - Staking operations Cost in Dollars. Cutting stakes and transporting 100.02 Staking 31*20 Tying 37.71 Cost of wrapping twine for tying 3*75 172.68 Pruning 35.61 35.61 Total expenses for staking and pruning operations 208,29 Total expenses for whole experiment 954*63 Cutlassing of the experimental beds was not costed for as the operation was given on contract to a local farmer who used the thrash to feed his Stock. Payment for the operation was therefore in kind. 65 A P P E N D I X I I I . Value of Produce For Different Treatments. Figures are derived from 1/19 of an acre and converted to cost per acre. Date of harvest­ Price per lb. F L A T M 0 U L D R I D G E ing. on day of harvest­ ing in cents. Yield/Acre in lbs Total valu« in $. BWT. Yield/Acre in lbs. Tbtal value in $ BWI. Yield/Acre in lbs. Total value in $ BWT. 4.2.58 50 361.00 180.50 256.50 128.25 399.00 199.50 7.2.58 45 408.50 183.53 593.75 267.19 864.50 389.03 11.2.58 25 1125.75 281.44 964.25 241.06 1349.00 337.25 14.2.58 25 1277.75 319.44 1410.75 352.69 1363.25 340.81 19.2.58 25 2370.25 592.56 3116.00 779.00 2455.75 613.94 21.2.58 30 1838.25 459.56 1529.50 382.38 1805.00 541.50 25.2.58 25 1890.50 472.63 1439.25 359.81 1985.50 496.38 27.2.58 25 969.00 242.25 1054.50 263.63 1187.50 296.88 3.3.58 26 931.00 242.06 1021.25 265.53 1230.25 319.87 6.3.58 20 346.75 69.35 631.75 126.35 722.00 144.40 10.3.58 20 370.50 74.10 593.75 118.75 308.75 61.75 Total 11889.25 3117.72 12611.25 3284.64 13670.50 3741.31 66 (B) Date of Price per lb. on STAKING & PRUNING STAKING NO PRUNING NO STA KINGj NO PRUNING harvest­ ing day of harvest­ ing in Cents. Yield/Acre in lbs. Total Value in $ BWI. Yield/Acre in lbs. Total Value in $ BWI Yield/Acre in lbs. Total value in $ BWI. 4.2.58 50 517.75 258.88 175.75 87.88 323.00 161.50 7.2.58 45 627.00 282.15 574.75 258.64 665.00 299.25 11.2.58 25 945.25 236.31 950.00 237.50 1543.75 385.94 14.2.58 25 1187.50 296.88 1263.50 315.88 1610.25 402.56 19.2.58 25 2479.50 619.88 3073.25 768.31 2389.25 597.31 21.2.58 30 1624.56 487.35 1800.25 540.07 1748.00 524.40 25.2.58 25 1306.25 326.56 2341.75 585.44 1667.25 416.81 27.2.58 25 1040.25 260.06 1021.25 255.31 1149.50 287.38 3.3.58 26 1467.75 381,62 983.25 255.65 722.00 187.72 6.3.58 20 505.50 100.70 722.00 144.40 475.00 95.00 10.3.58 20 769.50 153.90 475.00 95.00 28.50 5.70 Total 12468.75 3404.29 13380.75 3544.08 12321.50 3363.57 Average value of Crop for Different Treatments. Average price per lb. during season - 29 cents. Treatment Average value of crop in $ BWI. flat 344-9.88 Mould 3657.26 Ridge 3964.45 Staking and pruning 3615.94 Staking no pruning 3880.42 No staking, no pruning 3573.24 m 67 A P P E N D I X I V , COST - PROFIT ANALYSIS For Staking and Pruning Treatments, ITEM Staking & Pruning Staking, No Pruning No Staking, No Pruning. Total value of early crop (fruit produced per acre during the first 4 pickings) in $ BWI, 1074.22 899.90 1249.24 Total value of fruit produced during the experiment in $ BWI. 3516.94 3880.42 3573.24 Total cost of production per acre in $ BWI. 457.07 421.46 248.78 Net income at end of Season in $ B-JI, 3158.87 3458.96 3324.46 - 68 - B I B L I O G R A P H Y y Allerton, F. W. Anonymous Atkinson, J. D. et alj Babb, M.F. Baker, C.E. Barret, R.J.R. Blommestein, A.V. & E. F. Malan Boswell, V.R. Brasher, E. P. & K. C. Westover Butler, L. Campbell, J.S. Casseres, E.H. Tomato Growing, Faber, 1954. Agricultural Report, Antigua, 1949. Agricultural Report, St. KLtts, 1931. Agricultural Report, St. Vincent, 1911. Tomato Diseases and Pests in New Zealand and their control, 1949. Residual effect of forcing and hardening of tomatoes, cabbages, and cauliflower - U.S. Department. Agr. Tech. Bull. 760 - 1940. Early fruiting of tomatoes as induced by the use of soluble phosphates. Amer. Soc. Hort. Sci. Proc. 35: 668 - 672. 1937. Investigations on dry-season tomato production in Trinidad - I.C.T.A. Unpublished Dissertation. 1954/55. Tomato cultivation in the low veldf Fmg.» in S. Africa. 29: 287 - 294. 1954. Improvement and genetics of tomatoes, peppers and egg plant, - U.S. Dept. Agric. Year Book. pp. 176 - 206. 1937. Effect on yield of hardening the tomato plant, - Amer. Soc. Hort. Sci. Proc. 35: 686 - 689. 1937. Inheritance of fruit size in the tomato - Canadian Jour. Res. C., 19: 216-244. 1941. Report on the cultivation of vegetables in Montserrat. 1957. Effect of date of sowing, spacing and foliage trinming of plants in flats and on yield of tomatoes - Amer. Hort. Sci. Proc. 50f 285. 1947. 69 Clayton, E.E. Chenery, E.M. Chin, E.S.A. Christidis, B.G. Cobley, L.S. Cochran, W.G. & G. M. Cox Craft, C.C. & P. H. Heinze Currence, T.M. Deoner, M. T. et al. Doolittle, S.P. Edgerton, C.V. & C.C. Moreland : Edwards, W.H. Evans, H.J. & R. V. Troxler. The relation of Temperature to the Fusarium wilt of tomato, - Amer. Jour. Bot. 10: 77 - 87, 1923. Soils of Central Trinidad, 1952. Fertilizer trials on Tomatoes - West Indian Agric, Montserrat, 1932. The Importance of the shape of plots in field experimentation - Jour. Agric. Sci. 21: 14 - 37, 1931. An Introduction to the Botany of Tropical Crops, 1956. Experimental Designs, John Wiley & S ons, 1957. Physiological Studies of mature green tomatoes in Storage - Amer. Soc. Hort. Sci. Proc. 64: 343 - 350. 1954. The interactions between variety, spacing and staking of tomato plants. Amer. Soc. Hort. Sci. Proc. 39: 315 - 318. 1941. Pruning and training Tomatoes in the South. - U.S. Dept. Agric. Circ. 712. 1944. Tomato Diseases - U.S. Dept. Agr. Farmer's Bull. 1934. 1948. Tomato wilt - La. Agr. Expt. Stat. Bull. 175 . 1920. Tomato pests and their control Dept. of Agri. Ext. Circ. No. 22. Jamaica, 1948. Relation of calcium nutrition to the incidence of blossom-end rot in Tomatoes - Amer. Soc. Hort. sci. Proc. 61: 346 - 352, 1953. 70 - j Fenwich, J. S, Frazier, W.A. et al. Gilbert, J.C. & D. C. McGuire Investigations into Dry Season tomato cultivation in Trinidad, - I.C.T. A. unpublished Dissertation 1956/57, Seven new tomatoe varieties resistant to spotted wilt, fusarium wilt and Gray leaf spot - Uni, Hawaii Agri, Expt, Stat. Bull 103, 1950, Inheritance to severe root knot from .Meloidogyne incognita in commercial types of tomatoes - Amer. Soc. Hort. Sci. Proc. 68: 437 - 442. 1956. Grace, N. H. Halsey, L.H. et al. Hammer, K.C. et al. Hardy, F. & R.E.D. Baker Hawthorne, L.R. Hester, J.B, Physiologic curve of response to phytohormones by seeds, growing plants, cuttings and lower plant forms - Canadian Jour. Res. 15: 538 - 546. 1937. Containers for shipping Florida tomatoes. Florida Agric. Ext. Sci. Bull. No.560. 1955. Effects of light intensity, day length, temperature, and other environmental factors on the ascorbic acid content of tomatoes - Journ. Nutr. 29: 85 - 97, 1945. Report on a visit to the Colonial Development Corporation tomato project, Andros Is. Bahamas, 1952. Pruning unstaked tomatoes - Amer. Soc. Hort. Sci.- Proc 37: 930 - 934. 1939. Absorption of nutrients by the tomato plant at different stages of growth. Amer. Soc. Hort. Sci. Proc. 36: 729. 1938. Hester, J. B, A review of soil practices with some outstanding crops in 1940 - Report of Ten ton Tomato Club of New Jersey, 1940. 71 Hester, J.B. _ et al. The efficient use of nitrogen in tomato culture - Amer. Soc. Hort. Sci. Proc. 32: 308. 1941. Rate and amount of plant nutrients absorbed by various vegetables - Amer. Soc. Hort. Sci. Proc. 57; 249, 1951. Hieronymus, A.M. Spacing and training tomatoes - uni. Hawaii Agr. Ext. Serv. Rev. Circ. 35, 1949. Holmes, E. Hoffman, i.e. Jenkins, J.A. Jones, L.G. G.F. Warren. Era us, E.J. & H.R. Kraybill Landrau, P. G. Samuel. Larson, R.E. Tomatoes in the Canaries. Trop. Agr. 8: 111 - 113. 1931. Present cultural methods in growing spring green house tomatoes in Ohio - Annual Report Vegetable Grower's Association of America - 1938. The origin of the cultivated tomato. Econ. Bot. 2: 379-392. 1948. The efficienty of various methods of application of phosphorus for tomatoes. - Amer. Soc. Hort. Sci. Proc. 63; 309 - 319. 1954. Vegetation and reproduction with special reference to the tomato - Ore. Agr. Expt. Sta. Bull. 149, 1948. The use of various fertilizer treatments on the Plamar variety of tomatoes growing on a Coto clay - Jour. Agric. Univ. Puerto Rico 39: ' . 1955. The Fj combining ability of certain tomato varieties - Amer. Soc. Hort. Sci. Proc. 39'J . 313 - 314, 1941, Lawrence, W.J.C. Science 1 and the Glass House, Oliver and Boyd, 1950. Lindstrom, E.W. The inheritance of crate and related shapes of tomato fruit. - Jour. Agr. Res. 34*. 961 - —«• . 985, 1927. 72 Loomis, W. E. Studies in the transplanting of vegetable plants Cornell Uni. Agr. Ext. Memoir 87, 1925. Mac Arthur, J.W. Inherited characters in the tomato. I. Hie self pruning habit - Jounr. Heredity 23; p 395 396. 1932. Mach, W.B. et al. Maclinn, W.A. C. R. Fellers. Magruder, R. McCallan, E.A. McCue, G.A. Miller, J.R. MLnges, p. A. et al. Howell, W. Nylund, R. E. Fertilizer experiments V7ith tomatoes 1919 -39. Pennsylvania State College Agric. Exp. Stat. Bull. 393. 1940. Ascorbic acid (Vitamin C) in tomatoes and tomato products. - Mass. Agri. Exp. Stat. Bull. 254 1938. Hie effect of pruning and staking upon production of tomatoes. - Amer. Soc. Hort. Sci. Proc. 21: 270 - 273. 1924. 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